Holotype tracker for Philippine tiger beetles (Coleoptera: Cicindelidae)

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[Main] [Tiger beetles of PH]

This tracker is created to assist researchers who are interested in studying the holotypes of Philippine tiger beetle species. The following is not a full list, but shall be constantly updated. The information presented here are based only on the publications listed under the references.

The International Code of Zoological Nomenclature (ICZN) defines a "holotype" as "the single specimen upon which a new nominal species-group taxon is based in the original publication" (Chapter 16, Article 73.1, reference 8).

The Wildlife Export Certification issued by the DENR for specimens that are transported outside of the country states that: 

"Further, upon completion of the Research purposes, representatives of all the species collected, excluding holotypes may be retained on deposit with the [foreign university] and all remaining specimens including holotypes shall be returned to the Philippines and deposited at the National Museum of the Philippines or [local university] museum. The specimens retained shall remain the exclusive property of the Philippine Government and must be returned upon request of the Department of Environment and Natural Resources (DENR)." 

As a quick visual aid, the following color coding is applied:

  • (Blue) Deposited in the National Museum of the Philippines
  • (Green) Deposited in Philippine local universities / local private collection
  • (Gold) Deposited outside of the Philippines, and collected AFTER the enactment of RA 9147
  • (Purple) Deposited outside of the Philippines, but collected BEFORE the enactment of RA 9147

The Holotypes of some Philippine Tiger beetles (Coleoptera: Cicindelidae)
  1. Calomera cabigasi Cassola, 2011 [distribution]
    • "Holotype deposited in the Staatliche Naturwissenschaftliche Sammlungen Bayerns (Zoologische Staatsammlung, Munich, Germany; ZSM)" quoted verbatim (page 130, reference 5)
    • Place collected: "Misamis or., Gingoog City" quoted verbatim (page 130, reference 5)
    • Year collected: 2001 (page 130, reference 5)
  2. Calomera mindanaoensis (Cassola, 2000) [distribution]
    • Note: "The genus Calomera Motschulsky, 1862  was formerly often quoted in the entomological literature under the junior synonym Lophyridia Jeannel, 1946" quoted verbatim (page 129, reference 5)
    • "Nationaal Natuurhistorisch Museum of Leiden, Netherlands (RMNH)" quoted verbatim (pages 492 & 500, reference 4)
    • Place collected: "Mindanao (Davao Oriental): Boston, Mt Agtuuganon, Camp 55" quoted verbatim (page 500, reference 4)
    • Year collected: 1996 (page 500, reference 4)
  3. Cylindera (Conidera) mindoroana Zettel and Wiesner, 2018
    • " Holotype male, (coll. Wiesner, Wolfsburg, Germany, long term loan of Natural History Museum Stuttgart)..." quoted verbatim (page 2, reference 15)
    • Place collected:  "Philippines, Mindoro, Mt. Halcon" quoted verbatim )page 2, reference 15)
    • Year collected: 2010
  4. Cylindera (Cylinderina) rothschildi 
    • "Holotype male in MNHN" quoted verbatim (page 8, reference 11). "MNHN Muséum National d’Histoire Naturelle, Paris, France" (quoted verbatim (page 2, reference 11).
    • Place collected: no specific data presented in reference 11
    • Year collected: no specific data presented in reference 11 (Most probably collected in the late 1800s).
  5. Cylindera (Cylinderina) sierramadrensis Wiesner and Dheurle, 2018 [distribution]
    • "Holotype male in JWGC" quoted verbatim (page 5, reference 11). "JWGC Jürgen Wiesner Collection, Wolfsburg, Germany." (page 2 reference 11)
    • Place collected: "Aurora, Dingalan,... Sierra Madre" quoted verbatim (page 5, reference 11)
    • Year collected: 2018
  6. Cylindera (Cylinderina) vandenberghei Dheurle, 2016 [distribution]
    • "MNHN, Muséum national d’Histoire naturelle, Paris, France" quoted verbatim (page 37, reference 14)
    • Place collected: "Philippines, North Luzon Island, Cagayan Province, Santa Ana" quoted verbatim (page 37, reference 14)
    • Year collected: 2015 (page 37, reference 14)
  7. Cylindera (Ifasina) fabiocassola Weisner, 1989
    • "in Coll. mea" quoted verbatim (page 238, reference 7). Probably in author's collection.
    • Place collected: "N. Luzon, Mt. Province" quoted verbatim (page 238, reference 7)
    • Year collected: 1986 (page 238, reference 7)
  8. Cylindera (Ifasina) mouthiezi Dheurle, 2015
    • "déposé au MNHN, Paris." quoted verbatim (page 117, reference 9). Translation: deposited in MNHN, Paris. 
    • Place collected: "Province de Bukidnon, Cabanglasan" quoted verbatim (page 117, reference 9)
    • Year collected: 2014
  9. Heptodonta mindoroensis Cassola, 2000 [distribution]
    • "Nationaal Natuurhistorisch Museum of Leiden, Netherlands (RMNH)" quoted verbatim (pages 492 & 496, reference 4)
    • Place collected: "Calapan, Mt Halcon, Barrio Luyang" quoted verbatim (page 496, reference 4)
    • Year collected: 1991 (page 496, reference 4)
  10. Myriochila philippinensis (Mandl, 1956)
    • "deposited in his private collection in Vienna, Austria. Later the holotype was transferred to the Museum G. Frey in Tutzing near Munich, Germany, and today it is part of the Georg Frey Collection in the Naturhistorisches Museum Basel, Switzerland" quoted verbatim (page 585, reference 16)
    • Place collected: "...municipality in Abra Province in the northern part of Luzon Island, the Philippines." quoted verbatim (page 585, reference 16)
    • Year collected: 1950's probably (page 585, reference 16)
  11. Neocollyris (Heterocollyris) chaudoiri sciakyi Anichtchenko and Medina, 2019 [distribution]
    • "in coll. R.Sciaky (Italy)" quoted verbatim (pages 13 & 14, reference 3)
    • Place collected: "Zamboanga del N / Gutalac" quoted verbatim (page 14, reference 3)
    • Year collected: 2015 (page 14, reference 3)
  12. Neocollyris (Neocollyris) anichtchenkoi Medina & Mapile,2023
    • "(MMCP)...., to be deposited at PNM."..."MMCP - Milton Medica Collection, Tagum City, Phillippines; PNM - Philippine National Museum" quoted verbatim (page 2230, reference 17) 
    • Place collected: "PHILIPPINES: Luzon, Pangasinan, Calasiao, Brgy. Nalsian Centro" quoted verbatim (page 2230, reference 17)
    • Year collected: 2023 (page 2230, reference 17)
  13. Thopeutica (Thopeutica) anichtchenkoi Wiesner, 2015 [distribution]
    • "SMNS, Staatliches Museum für Naturkunde, Stuttgart, Germany (at present in coll. WIESNER)." quoted verbatim (page 2, reference 12)
    • Place collected: "Mt. Apo" quoted verbatim (page 2, reference 12)
    • Year collected: 2011 (page 2, reference 12)
  14. Thopeutica (Thopeutica) barsevskisi Medina, Cabras, and Villanueva, 2020
    • "The holotype of this new species is deposited in the University of Mindanao Coleoptera Research Center (UMCRC)" quoted verbatim (page 95, reference 1)
    • Place collected: "Bohol, / Antiquera" quoted verbatim (page 96, reference 1)
    • Year collected: 2019 (page 96, reference 1)
  15. Thopeutica (Thopeutica) hinabanganensis Weisner, 2015 [distribution]
    • "SMNS, Staatliches Museum für Naturkunde, Stuttgart, Germany (at present in coll. WIESNER)" quoted verbatim (page 5, reference 12)
    • Place collected: "Central Visayas, Samar, Hinabangan" quoted verbatim (page 5, reference 12). Note: Samar belongs to Region VIII - Eastern Visayas, not Region VII - Central Visayas. The provinces of Central Visayas are Bohol, Cebu, Negros Oriental, and Siquijor.
    • Year collected: 2014 (page 5, reference 12)
  16. Thopeutica (Thopeutica) milanae Wiesner, 1992
    • "Holotypus und zwei Paratypen im Naturkundemuseum Stuttgart" quoted verbatim (page 128, reference 6). Translation: Holotype and two paratypes in the Stuttgart Natural History Museum
    • Place collected: "Visca N Baybay, Primärwald" quoted verbatim (page 129, reference 7). Translation: Visayas State College of Agriculture (now Visayas State University) Province of Leyte, Baybay City, Primary forest.
    • Year collected: 1991 (page 128, reference 6)
  17. Thopeutica (Thopeutica) pangantihoni Cassola and Zettel, 2006 [distribution]
    • "...holotype, allotype, and some paratypes in the University of San Carlos (Biological Collections), Cebu City, Philippines" quoted verbatim (page 46, reference 13)
    • Place collected: "Philippines, Quezon Province, Polillo Island, Panukulan, water shed area west of town." quoted verbatim (page 46, reference 13)
    • Year collected: 2004 (page 46, reference 13)
  18. Thopeutica (Thopeutica) petertaylori Medina, Cabras and Wiesner, 2019 [distribution]
    • "Holotype in UMCRC" quoted verbatim (page 2, reference 10). "UMCRC  University of Mindanao, Coleoptera Research Center, Davao City, Philippines" (page 1, reference 10)
    • Place collected: "Compostella Valley,/ New Bataan , Cagan,/" quoted verbatim (page 2, reference 10)
    • Year collected: 2012
  19. Thopeutica (Thopeutica) rolandmuelleri Cassola, 2000 [distribution]
    • "Nationaal Natuurhistorisch Museum of Leiden, Netherlands (RMNH)" quoted verbatim (pages 492 & 504, reference 4)
    • Place collected: "Mindanao (Davao Oriental): Boston, Mt. Agtuuganon, Camp 55" quoted verbatim (page 504, reference 4)
    • Year collected: 1996 (page 504, reference 4)
  20. Thopeutica (Thopeutica) wiesneri Anichtchenko and Medina, 2020 [distribution]
    • "DUBC- Daugavpils University Beetles collection (Latvia)" quoted verbatim (pages 389 & 391,  reference 2)
    • Place collected: "Mindanao, New Bataan, Cagan" quoted verbatim (page 391, reference 2)
    • Year collected: 2019 (page 391, reference 2)

References:

[1] Medina M. N. D., Cabras A. A., Villanueva R. J. T. 2020. Thopeutica (Thopeutica) barsevskisi sp. n. a new tiger beetle (Coleoptera: Cicindelidae) in Bohol Island Philippines. Baltic J. Coleopterol., 20 (1): 95 – 100.

[2] Anichtchenko, A. and Medina, M.M. 2020. Contribution to the knowledge of three species of the genus Thopeutica Schaum, 1861 from the Philippines (Coleoptera: Carabidae: Cicindelidae). Zootaxa. Vol. 4748(2): 389-395.

[3] Anichtchenko, A. and Medina, M.N. 2019. A New Neocollyris (Heterocollyris) Subspecies from Mindanao, Philippines (Coleoptera, Carabidae, Cicindelinae). Acta Biol. Univ. Daugavpils. Vol. 19(1); 13-15.

[4] Cassola, F.  2000. Studies on Tiger Beetles. CII. The Cicindelidae Collected by Roland Müller in the Philippine Islands, with Description of Three New Species (Coleoptera: Cicindelidae). 
Zool. Med. Leiden. Vol. 73; 491-509. 

[5] Cassola, F. 2011. Studies of Tiger Beetles. CLXXXIX. A new Calomera Species from Mindanao, Philippines. Zoologische Staatssammlung Munchen/Verlag Friedrich Pfeil. Vol.34(1).

[6] Wiesner, J. 1992. Eine neu Thopeutica von den Philippinen (Coleoptera: Cicindelidae), 26, Beitrag zur Kenntnis der Cicindelidae. Pages 128-130.

[7] Weisner, J. 1989. Beiträge zur Kenntnis der Philippinischen Cicindelidae (II), (Coleoptera), 22. Beitrag zur Kenntnis der Cicindelidae.Entomologische Zeitschrift 99, 16, 237 - 238.

[8] International Code of Zoological Nomenclature. Chapter 16, Article 73.1. Holotypes. https://code.iczn.org

[9] Dheurle, C. 2015. Cylindera (Ifasina) mouthiezi, nouvelle espèce des Philippines (Coleoptera Cicindelidae). L’Entomologiste, tome. 71(2): 117-118.

[10] Medina, M.N. D., Cabras, A.A., and Wiesner J. 2019. Thopeutica petertaylori, a new tiger beetle species (Colpeoptera: Cicindelidae) from Mindanao, Philippines. Insecta Mundi: A Journal of world Insect Systematics. 0733; 1-5.

[11] Wiesner J. and Dheurle C. 2018. Remarks on the subgenus Cylinderina Rivalier (Coleoptera: Cicindelidae: Cylindera) from the Philippines with description of a new species. Insecta Mundi. 0670; 1-16.

[12] Wiesner J. 2015. Two new Thopeutica species from the Philippines. Mitteilungen des Internationalen Entomologischen Vol. 40; 1-8.

[13] Cassola, F., Zettel, H. 2006. A new species and a new record of Thopeutica Chaudoir, 1861 from Polillo Island, Quezon Province, the Philippines (Coleoptera: Cicindelidae). Z. Arb. Ge, Ost.Ent. 58; 45-52.

[14] Dheurle, C. 2016. Une nouvelle espece de Cylindera Westwood, 1831, des Philippines (Coleoptera, Cicindelidae). Bulletin de la Societe entomologique de France. 121(1); 37-40.

[15] Zettel H. and Wiesner J. 2018. Cylindera (Conidera) mindoroana sp. n. (Coleoptera: Cicindelidae), A New Tiger Beetle Species from the Philippines. Insecta Mundi. Vol. 0632; 1-10.

[16] Zettel, H. and Pangantihon, C. V. 2015. On the Identity of the Tiger Beetle Cicindela philippinensis Mandl, 1956 (Coleoptera: Carabidae: Cicindelinae). Zootaxa 3941 (4); 585-588.

[17] Medina, M.N.D. & Mapile, D.O. 2023. Description of a New Species of Neocollyris W. Horn, 1901(Coleoptera: Cicindelidae) with a Detailed Description of the Larval Development Observed In Situon a Native Non-endemic Premna odorata Blanco(Lamiaceae) in the Philippines. Philippine Journal of Science.152 (6A): 2229-2237




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